Biodiversity Taxonomy Animalia Arthropoda Insecta Porifera Demospongiae Chordata Coleoptera Chrysomelidae Gulf of Mexico Amphibia Anura Aromobatidae Allobates Mexico Campeche Bank Malacostraca Mesomphalia Hemiptera Haplosclerida Anthophila Apoidea Marine Biology Yucatan fishes orchid bees pollinators Actinopterygii Caribbean Coral Hadziidae Mexican Atlantic Perciformes morphology reef species richness Decapoda Mammalia SEM Serranidae amphipods decapods groundwater isopods mysids sinkholes stygiomysids thermosbaenaceans Yucatan Peninsula cenote description endemic mitochondrial marker sinkhole subterranean Cicadidae Mollusca